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Both flies and mosquitoes pollinate flowers to some extent. In places where bees are less prevalent, this may become significant. BTW, there are a lot of different species of fly, some more obviously beneficial than others.
Try googling "flies pollinate" and "mosquitoes pollinate" for some interesting links.
Also, both flies and mosquitoes serve as food for other species, such as fish, birds, bats, and other insects.
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