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You can overclock a Harpertown. You can throw two Harpertowns in there and overclock them both, but you'll need to upgrade the fan (which you can do on a Mac).
The Mac Pro takes up to 32GB RAM (8x4GB). It has 4 hard drive bays, but you can fit more in there. I saw a photo of one with like 8 or 9 hard drives. Can you imagine 9TB? Or better still get 1.2TB on SAS at 15k rpm. So long as you've got the RAID card, you'll be fine. Want decent video? It's ready for 4 cards. It can be shipped with the NVIDIA Quadro FX 5600 1.5GB.
It seems perfectly customizable. A real pro could probably mod the iMac quite a bit, too.
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