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My wife wears thongs all the time and finds them extremely comfortable. I would suggest that it has to do with 1) fit and 2) material. My wife's are mostly really good cotton, and they fit perfectly.
I have worn a thong on several occassions myself and find that they are comfortable, just not the same as underwear that does not live up your crack. I thikn it has something to do with getting used to them. I find them a little naughty, which gives me an almost permanent erection, which makes them slightly tighter, which is sometimes not so comfortable. However, I doubt that your girlfriend will have the same problem, but I don't you you very well so...
Pierre
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